Fall back on `In-Reply-To` header for reply by email when sub-addressing with the reply key is not available.
See the discussion starting at https://gitlab.com/gitlab-org/gitlab-ce/issues/1360#note_1970380.
We could add the key to References, and read it from In-Reply-To as a fallback. Good idea.
Note that we're planning to add a feature to create an issue by email at some point, which will only work with a project/user-specific key in the address.
We need to expand the docs on the different options (sub-addressing, catch-all box, In-Reply-To).